Chalicodein Sapotherium

Chalicodein's arms were long and heavily clawed, adapted for browsing but forcing them to walk on their knuckles only.

The arms were used to reach for the branches of large trees and bring them close to its short trunk and reversed tusks to strip them clean of leaves. The way it used its curious tusks has been much debated. It may have rooted in soil for underground plant parts like roots and tubers, stripped branches for softer leaves, or stripped soft bark from tree trunks. This may have left its hands available for tool use.

Callosities on the ischium imply that these animals would sit on their haunches for extended periods of time, probably while feeding. Pad-supporting bony growth on the dorsal side of the manual phalanges is interpreted as evidence of knuckle-walking, which would probably be useful to avoid wearing down the claws, preserving them for use either as a forage-collecting rake or as a formidable defensive weapon, or both.

All of these characteristics show some convergence with other creatures such as elephants, ground sloths, great apes, bears (especially giant pandas), and a group of theropod dinosaurs known as therizinosaurs.